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Hamano, Tatsuo / Professor (by-work)   [Detail]
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Year of Birth: 1959 A.D.
Phone: +81 - 88 - 683 - 7027 E-mail:  h a m a n o . t a t s u o (@) t o k u s h i m a - u . a c . j p 
Academic Degree: Doctor of Agriculture Field of Study: stock-enhancement, 地域生物応用学, aquaculture
Lecture: Agritechnological Science Ⅱ (Bachelor Course), オリエンテーション (Graduate School), Special Lecture on Aqua-Field Biology (Graduate School), Field and Environmental Ecology (Bachelor Course), Project Research II (Graduate School), Undergraduate Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Advansed Exercise on Regional Science II (Graduate School), Aquatic Bioproduction Science (Bachelor Course), Fisheries Resources Biology (Bachelor Course), Advanced Theory for Environmental Management (L) (Graduate School), Advanced Theory for Environmental Management (S) (Graduate School), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Methods for Environmental Symbiosis Research (Graduate School), Environmental Science (Graduate School), Ecology Ⅱ (Bachelor Course), Practice for Agriscience A (Bachelor Course), Practice for Agriscience C (Bachelor Course), Practice for Agriscience I (Bachelor Course), Practice for Agriscience II (Bachelor Course), Field Practice for Agriculture, Forestry and Fisheries (Bachelor Course), Special Practice on Agrobioscience (Graduate School), Advanced Exercise on Agrobioscience (Graduate School), Advanced Research on Agrobioscience (Graduate School), Special Lecture on Agrobioscience (Graduate School), Research on Bioresource Science (Graduate School), Bio-Resource and Environmental Science (Bachelor Course), Bioindustry C (Bachelor Course), Practice for Bioscience and Bioindustry (Bachelor Course), English for Bioscience and Bioindustry (Bachelor Course), Basic bioindustry (Bachelor Course), Introduction to Field Science for Production (Bachelor Course), 研究計画発表会・博士論文中間発表会(総合科学課題研究Ⅱ) (Graduate School), themed Research in integrated Sciences 2 (Graduate School), Reading Scientific Papers (Bachelor Course), Reading Scientific Papers Ⅰ (Bachelor Course), Reading Scientific Papers Ⅱ (Bachelor Course), Basic Agroscience (Bachelor Course)
Subject of Study: 沿岸や河川の生物,希少生物の野生資源の増殖技術開発 (stock-enhancement, conservation, fishway, fish reef), 地域の特産的資源である生物について総合的に研究する (stock-enhancement, conservation), aquabiology (aquaculture, river, conservation)
Hattori, Takefumi / Professor (by-work)   [Detail]
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Year of Birth: 1962 A.D.
Phone: +81 - 88 - 656 - 7183 E-mail:  t h a t t o r i (@) i a s . t o k u s h i m a - u . a c . j p 
Academic Degree: 京都大学農学博士 Field of Study: Metabolic science for forest plants and microorganisms
Lecture: Agritechnological Science Ⅰ (Bachelor Course), Conservation Biology (Bachelor Course), Undergraduate Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Advansed Exercise on Regional Science II (Graduate School), Metabolic Science for Woody Plants and Forest Microorganisms (Bachelor Course), Advanced seminar in metabolism for forest plants and microorganisms (Graduate School), Forest biology (Graduate School), Forest Science (Bachelor Course), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Methods for Environmental Symbiosis Research (Graduate School), Practice for Agriscience A (Bachelor Course), Practice for Agriscience C (Bachelor Course), Practice for Agriscience I (Bachelor Course), Practice for Agriscience II (Bachelor Course), Field Practice for Agriculture, Forestry and Fisheries (Bachelor Course), Advanced Exercise on Agrobioscience (Graduate School), Advanced Research on Agrobioscience (Graduate School), Advanced course in metabolic science for couservation of forest ecosystem (Graduate School), Advanced seminar in metabolic science for conservation of forest ecosystem (Graduate School), Research on Bioresource Science (Graduate School), Bioindustry C (Bachelor Course), Practice for Bioscience and Bioindustry (Bachelor Course), English for Bioscience and Bioindustry (Bachelor Course), Basic bioindustry (Bachelor Course), Introduction to Field Science for Production (Bachelor Course), Developed environmental symbiosis studies (Graduate School), Reading Scientific Papers (Bachelor Course), Reading Scientific Papers Ⅰ (Bachelor Course), Reading Scientific Papers Ⅱ (Bachelor Course)
Subject of Study: Elucidation of carbon metabolism of wood-rotting fungi and ectomycorrhizal fungi, Development of utilization method for forest products, Biosynthesis and biodegradation of secodary metabolites in plants (wood-rotting fungi, ectomycorrhizal fungi, forest products, forest science, oxalic acid)
Imai, Shoji / Professor (by-work)   [Detail]
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Year of Birth: 1961 A.D.
Phone: +81 - 88 - 656 - 7273 E-mail:  s h o j i . i m a i (@) t o k u s h i m a - u . a c . j p 
Academic Degree: Doctor of Science Field of Study: Analytical Chemistry, Environmental Analytical Chemistry, Analytical Atomic Spectrometry, Environmental Chemistry
Lecture: Project Research I (Graduate School), Analytical Chemistry 1 (Bachelor Course), Analytical Chemistry 2 (Bachelor Course), Analytical Chemistry I (Bachelor Course), Introductory Laboratory for Chemistry (Bachelor Course), Laboratory for Chemistry 1 (Bachelor Course), Laboratory for Chemistry 2 (Bachelor Course), Experiments in Chemistry I (Bachelor Course), Experiments in Chemistry II (Bachelor Course), Graduation Work (Bachelor Course), Regional Science II (Graduate School), Advanced Exercise on Regional Science I (Graduate School), Advansed Exercise on Regional Science II (Graduate School), Outline of Chemistry (Common Curriculum), Seminar on Natural Sciences (Bachelor Course), Technical English Fundamentals 2 (Bachelor Course), Inorganic Chemistry II (Bachelor Course), Advanced Material Chemistry (Graduate School), Seminar in Physics, Chemistry, and Earth Science (Bachelor Course), Introduction to Science and Technology (Common Curriculum), Interdisciplinary Seminar in Science and Technology (Graduate School), Adovanced Evironmental System Analysis (Graduate School), Experiments in Symbiotic Environmental Studies II (Bachelor Course), Advanced Environmental Analytical Chemistry (Graduate School), Instrumental Analytical Chemistry for Environment (Bachelor Course), Advanced Environmental Inorganic Chemistry (Graduate School), Graduate Research on Natural Sciences (Graduate School), Graduate Seminar on Natural Sciences (Graduate School), Reading Scientific Papers (Bachelor Course)
Subject of Study: Toxic Trace elemet analysis (environmental analytical chemistry, environmental chemistry, environmental science), Toxic trace elemet analysis in environments, Atomization mechanism of Lead, Gold, Arsenic, Indium in GFAAS, 環境分析化学 (environmental chemistry, environmental analytical chemistry, natural environment)
Kanemaru, Kaori / Professor (by-work)   [Detail]
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Year of Birth: 1958 A.D.
Phone: +81 - 88 - 656 - 7268 E-mail:  k a n e m a r u (@) t o k u s h i m a - u . a c . j p 
Academic Degree: Doctor of Philosophy Field of Study: Food Processing and Preservation Science, Functional Food Science, Cookery Science
Lecture: SIH(Strike while the Iron is Hot) Training (Common Curriculum), Food Science (Bachelor Course), Undergraduate Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Basic Food Chemistry (Bachelor Course), Methods for Environmental Symbiosis Research (Graduate School), Advanced Bioresource Chemistry (Graduate School), Research on Bioresource Science (Graduate School), Bioindustry B (Bachelor Course), Practice for Bioscience and Bioindustry (Bachelor Course), English for Bioscience and Bioindustry (Bachelor Course), Basic bioindustry (Bachelor Course), Special lecture on bioscience and bioindustry (Bachelor Course), Reading Scientific Papers (Bachelor Course), Reading Scientific Papers Ⅰ (Bachelor Course), Reading Scientific Papers Ⅱ (Bachelor Course), Environmental and Natural Resource Economics I (Graduate School), Environmental and Natural Resource Economics II (Graduate School), Enzymology (Bachelor Course), Introduction to Food and Health (Bachelor Course), Food and Bioresource act (Bachelor Course), Food Processing and Preservation (Bachelor Course), Advanced Lecture on Food Processing Preservation Science (Graduate School), Advanced Lecture of Food Safety Science (Graduate School), Food Engineering (Bachelor Course), Food Microbiology (Bachelor Course), Food Hygienics (Bachelor Course), Food Hygienics I (Bachelor Course), Advanced Lecture on Food Safety Science (Graduate School), Special Practice on Food Bioscience (Graduate School), Advanced Exercise on Food Bioscience (Graduate School), Advanced Research on Food Bioscience (Graduate School), Special Lecture on Food Bioscience (Graduate School), Basic practice for food science (Bachelor Course), Practice for food science A (Bachelor Course), Practice for food science B (Bachelor Course), Practice for food science C (Bachelor Course), Exercise in food science Ⅰ (Bachelor Course), Exercise in food science Ⅱ (Bachelor Course)
Subject of Study: 食品成分による抗微生物作用と食品保蔵への応用 (食中毒菌, 腐敗菌, 抗菌活性, 抗酸化活性, 食品加工), 食品成分や環境物質による細胞への影響 (細胞保護作用, 細胞毒性, 抗ガン活性, 有用菌増殖活性, 免疫賦活活性, 食品資源の循環利用), 調理文化の地域性と郷土食の調査研究 (郷土食, 食文化, 調理)
Makabe, Kazuhiro W. / Professor (by-work)   [Detail]
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Year of Birth:
Phone: E-mail:
Academic Degree: Doctor of Science Field of Study: Molecular and Evolutionary Developmental Biology
Lecture: STEM Practice (Bachelor Course), Genomic and Genetic Technology (Common Curriculum), Advanced Biotechnology (Bachelor Course), Molecular Develomental Biology (Bachelor Course), Graduation Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Advansed Exercise on Regional Science II (Graduate School), Seminar on Natural Sciences (Bachelor Course), Technical English Fundamentals 2 (Bachelor Course), Interdisciplinary Seminar in Science and Technology (Graduate School), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Bioscience and Technology (Bachelor Course), Introductory Laboratory for Bioscience (Bachelor Course), Laboratory for Bioscience 1 (Bachelor Course), Laboratory for Bioscience 3 (Bachelor Course), Developmental Biology (Bachelor Course), Advanced Developmental Bioinformatics (Graduate School), Developmental Genetics (Bachelor Course), Cell Engineering (Bachelor Course), Graduate Research on Natural Sciences (Graduate School), Graduate Seminar on Natural Sciences (Graduate School), Reading Scientific Papers (Bachelor Course)
Subject of Study: ホヤを用いた初期発生メカニズムの分子·進化発生学的研究 (ホヤ, 母性因子, 局在RNA, genome, EST, 転写因子ネットワーク, 小分子RNA)
Matsuo, Yoshinori / Professor (by-work)   [Detail]
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Year of Birth: 1959 A.D.
Phone: 656 - 7270 E-mail:  m a t s u o (@) i a s . t o k u s h i m a - u . a c . j p 
Academic Degree: Doctor of Science Field of Study: Genetics
Lecture: Graduation Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Advansed Exercise on Regional Science II (Graduate School), Basic Biology (Common Curriculum), Seminar on Natural Sciences (Bachelor Course), Technical English Fundamentals 1 (Bachelor Course), Technical English Fundamentals 2 (Bachelor Course), Interdisciplinary Seminar in Science and Technology (Graduate School), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Methods for Environmental Symbiosis Research (Graduate School), Advanced Lecture of Adaptation for Environment (Graduate School), Bioscience and Technology (Bachelor Course), Introductory Laboratory for Bioscience (Bachelor Course), Laboratory for Bioscience 1 (Bachelor Course), Laboratory for Bioscience 3 (Bachelor Course), Biostatistics (Bachelor Course), Research Approaches in Science and Technology D (Graduate School), Graduate Research on Natural Sciences (Graduate School), Graduate Seminar on Natural Sciences (Graduate School), Adaptive Evolutionary Biology (Bachelor Course), Genetics and Evolution (Common Curriculum), Advanced lecture of genetic science for environment (Graduate School), Population Genetics (Bachelor Course), Population Genetics (Graduate School), Reading Scientific Papers (Bachelor Course)
Subject of Study: Genetic mechanism of adaptive evolution (Drosophila, Mechanism of adaptive evolution, Gene family, Concerted evolution, GC contents)
Watanabe, Minoru / Professor (by-work)   [Detail]
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Year of Birth: 1963 A.D.
Phone: +81 - 88 - 656 - 7253 E-mail:  m i n o r u (@) t o k u s h i m a - u . a c . j p 
Academic Degree: Doctor of Science Field of Study: Molecular Biology, Developmental Biology
Lecture: SIH(Strike while the Iron is Hot) Training (Common Curriculum), Advanced Biotechnology (Bachelor Course), Molecular Biology (Bachelor Course), Graduation Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Basic Biology (Common Curriculum), Basic Biology Experiments (Common Curriculum), Basic Biology Experiments (Common Curriculum), Seminar on Natural Sciences (Bachelor Course), Technical English Fundamentals 2 (Bachelor Course), Interdisciplinary Seminar in Science and Technology (Graduate School), Methodology of Teaching Science I (Bachelor Course), Methodology of Teaching Science II (Bachelor Course), Methodology of Teaching Science III (Bachelor Course), Methodology of Teaching Science IV (Bachelor Course), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Advanced Biosciences (Graduate School), Bioscience and Technology (Bachelor Course), Life science and modern society (Common Curriculum), Introductory Laboratory for Bioscience (Bachelor Course), Laboratory for Bioscience 2 (Bachelor Course), Laboratory for Bioscience 3 (Bachelor Course), Biology -Fundamental Cell Biology- (Common Curriculum), Advanced Lecture of Developmental Bioinformatics (Graduate School), Functional Cell Biology (Bachelor Course), Graduate Research on Natural Sciences (Graduate School), Graduate Seminar on Natural Sciences (Graduate School), Genetic Engineering (Bachelor Course), Advanced lecture of genetic science for environment (Graduate School), Reading Scientific Papers (Bachelor Course)
Subject of Study: Molecular biological study of the early frog development (Xenopus laevis, Transcriptional Regulation, signal transduction, RNAi, development, morphogenesis, transcription factor, genome editing)
Kawakami, Ryushi / Associate Professor (by-work)   [Detail]
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Year of Birth: 1977 A.D.
Phone: +81 - 88 - 656 - 7247 E-mail:  k a w a k a m i (@) t o k u s h i m a - u . a c . j p 
Academic Degree: Ph.D Field of Study: Enzyme Chemistry
Lecture: Food Science (Bachelor Course), Undergraduate Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Advanced lecture of environmental symbiotics (Graduate School), Biochemistry 3 (Bachelor Course), Research on Bioresource Science (Graduate School), Advanced lecture of Bioresources (Graduate School), Bioindustry B (Bachelor Course), Practice for Bioscience and Bioindustry (Bachelor Course), English for Bioscience and Bioindustry (Bachelor Course), Reading Scientific Papers (Bachelor Course), Reading Scientific Papers Ⅰ (Bachelor Course), Reading Scientific Papers Ⅱ (Bachelor Course), Genetic Engineering (Bachelor Course), Biotechnology of Genetic Resources (Bachelor Course), Enzymology (Bachelor Course), Advenced enzyme chemistry (Graduate School), Introduction to Food and Health (Bachelor Course), Advanced Exercise on Food Bioscience (Graduate School), Advanced Research on Food Bioscience (Graduate School), Basic practice for food science (Bachelor Course), Practice for food science A (Bachelor Course), Practice for food science B (Bachelor Course), Practice for food science C (Bachelor Course), Exercise in food science Ⅰ (Bachelor Course), Exercise in food science Ⅱ (Bachelor Course)
Subject of Study: 極限環境微生物酵素の機能・構造・応用 (超好熱アーキア, アミノ酸脱水素酵素, アミノ酸ラセマーゼ, タンパク質結晶化, X線結晶構造解析)
Satoh, Masaya / Associate Professor (by-work)   [Detail]
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Year of Birth: 1962 A.D.
Phone: +81 - 88 - 656 - 7222 E-mail:  s a t o h . m a s a y a (@) t o k u s h i m a - u . a c . j p 
Academic Degree: PhD Field of Study: Biology
Lecture: SIH(Strike while the Iron is Hot) Training (Common Curriculum), Talk about your favorites (Common Curriculum), Career Paths (Bachelor Course), Undergraduate Work (Bachelor Course), Regional Science I (Graduate School), Advanced Exercise on Regional Science I (Graduate School), Forest biology (Graduate School), Advanced Lecture on Plant Protection Science (Graduate School), Introduction to Botany (Bachelor Course), Plant Environmental Stress Physiology (Bachelor Course), Plant Pathology (Bachelor Course), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Advanced lecture of environmental pollutants (Graduate School), Environmental Physiology (Bachelor Course), Practice for Agriscience A (Bachelor Course), Practice for Agriscience C (Bachelor Course), Practice for Agriscience I (Bachelor Course), Practice for Agriscience II (Bachelor Course), Field Practice for Agriculture, Forestry and Fisheries (Bachelor Course), Advanced Exercise on Agrobioscience (Graduate School), Advanced Research on Agrobioscience (Graduate School), Research on Bioresource Science (Graduate School), Bio-Resource and Environmental Science (Bachelor Course), Bioindustry C (Bachelor Course), Practice for Bioscience and Bioindustry (Bachelor Course), English for Bioscience and Bioindustry (Bachelor Course), Introduction to Field Science for Production (Bachelor Course), Civil and Envioronmental Studies (Bachelor Course), Project for Integrated Arts and Sciences (Bachelor Course), Nature Conservation (Common Curriculum), Reading Scientific Papers (Bachelor Course), Reading Scientific Papers Ⅰ (Bachelor Course), Reading Scientific Papers Ⅱ (Bachelor Course)
Subject of Study: Introduction and propagation of Ginkgo biloba in Japan, Mechanism of detoxification in phytoplankton, Genetic diagnosis for serious Strawberry diseases, History of relationship between giant trees and human (history)
Yamamoto, Takashi / Associate Professor (by-work)   [Detail]
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Year of Birth: 1971 A.D.
Phone: +81 - 88 - 656 - 7263 E-mail:  t a k a s h i - y a m a m o t o . i a s (@) t o k u s h i m a - u . a c . j p 
Academic Degree: 博士(工学) Field of Study: Environmental Physical Chemistry, Catalytic Chemistry, X-ray Spectrometry
Lecture: SIH(Strike while the Iron is Hot) Training (Common Curriculum), オリエンテーション1年 (Bachelor Course), Environmental Physical Chemistry (Graduate School), Analytical Chemistry 1 (Bachelor Course), Analytical Chemistry 2 (Bachelor Course), Introductory Laboratory for Chemistry (Bachelor Course), Laboratory for Chemistry 1 (Bachelor Course), Laboratory for Chemistry 2 (Bachelor Course), Experiments in Chemistry I (Bachelor Course), Experiments in Chemistry II (Bachelor Course), Graduation Work (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Advansed Exercise on Regional Science II (Graduate School), Basic Chemistry Experiments (Common Curriculum), Seminar on Natural Sciences (Bachelor Course), Technical English Fundamentals 2 (Bachelor Course), Inorganic Chemistry 1 (Bachelor Course), Inorganic Chemistry 2 (Bachelor Course), Inorganic Chemistry I (Bachelor Course), Inorganic Chemistry II (Bachelor Course), Physical Chemistry 1 (Bachelor Course), Physical Chemistry 2 (Bachelor Course), Physical Chemistry I (Bachelor Course), Physical Chemistry II (Bachelor Course), Advanced Material Chemistry (Graduate School), Environmental Physical Chemistry (Graduate School), Seminar in Physics, Chemistry, and Earth Science (Bachelor Course), Interdisciplinary Seminar in Science and Technology (Graduate School), Experiments in Symbiotic Environmental Studies II (Bachelor Course), Advanced Environmental Physical Chemistry (Graduate School), Graduate Research on Natural Sciences (Graduate School), Graduate Seminar on Natural Sciences (Graduate School), Reading Scientific Papers (Bachelor Course)
Subject of Study: 新規固体触媒の開発と物性発現機構解明, X線分析化学 (固体酸塩基, 複合酸化物, ナノレベル構造解析, XAFS, 環境触媒)
Yamashiro, Tadashi / Associate Professor (by-work)   [Detail]
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Year of Birth: 1972 A.D.
Phone: +81 - 88 - 656 - 7257 E-mail:  t y a m a s h (@) i a s . t o k u s h i m a - u . a c . j p 
Academic Degree: Dr. of Science Field of Study: Biology, 植物系統分類学
Lecture: Global Communication - Traditional art craft - (Common Curriculum), Molecular Ecology (Graduate School), Undergraduate Work (Bachelor Course), Lecture on Museum Materials (Bachelor Course), Advanced Exercise on Regional Science I (Graduate School), Forest biology (Graduate School), Introduction to Botany (Bachelor Course), Experiments in Symbiotic Environmental Studies III (Bachelor Course), Advanced Lecture of Environmental Symbiotic Biology (Graduate School), Advanced Lecture of Adaptation for Environment (Graduate School), Introduction to Biodiversity (Bachelor Course), Practice for Agriscience A (Bachelor Course), Practice for Agriscience C (Bachelor Course), Practice for Agriscience I (Bachelor Course), Practice for Agriscience II (Bachelor Course), Field Practice for Agriculture, Forestry and Fisheries (Bachelor Course), Advanced Exercise on Agrobioscience (Graduate School), Advanced Research on Agrobioscience (Graduate School), Research on Bioresource Science (Graduate School), Bio-Resource and Environmental Science (Bachelor Course), Bioindustry C (Bachelor Course), Practice for Bioscience and Bioindustry (Bachelor Course), English for Bioscience and Bioindustry (Bachelor Course), Introduction to Field Science for Production (Bachelor Course), Phylogeny and Taxonmy of Plants (Bachelor Course), Nature Conservation (Bachelor Course), Reading Scientific Papers (Bachelor Course), Reading Scientific Papers Ⅰ (Bachelor Course), Reading Scientific Papers Ⅱ (Bachelor Course)
Subject of Study: 被子植物の分類・系統
Nakamura, Mitsuhiro / Associate Professor (by-work)   [Detail]
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Year of Birth: 1974 A.D.
Phone: +81 - 88 - 656 - 7246 E-mail:  n a k a m u r a . m i t s u h i r o (@) t o k u s h i m a - u . a c . j p 
Academic Degree: 農学博士 Field of Study: Chemistry, Natural Products Chemistry, Bioorganic Chemistry
Lecture: STEM Practice (Bachelor Course), Introductory Laboratory for Chemistry (Bachelor Course), Laboratory for Chemistry 1 (Bachelor Course), Laboratory for Chemistry 2 (Bachelor Course), Experiments in Chemistry I (Bachelor Course), Experiments in Chemistry II (Bachelor Course), Graduation Work (Bachelor Course), Basic Bio-organic Chemistry (Common Curriculum), Basic Chemistry Experiments (Common Curriculum), Seminar on Natural Sciences (Bachelor Course), Technical English Fundamentals 2 (Bachelor Course), Spectrometric Identification of Organic Compounds (Bachelor Course), Organic Functional Material Chemistry (Graduate School), Advanced Material Chemistry (Graduate School), Seminar in Physics, Chemistry, and Earth Science (Bachelor Course), Interdisciplinary Seminar in Science and Technology (Graduate School), Experiments in Symbiotic Environmental Studies II (Bachelor Course), Biological and Organic Chemistry (Bachelor Course), Graduate Research on Natural Sciences (Graduate School), Graduate Seminar on Natural Sciences (Graduate School), Reading Scientific Papers (Bachelor Course)
Subject of Study: 機能性天然物質の探索と高機能化 (機能性物質, 生物発光基質, 硫酸化)